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On the cover. Brachypodium distachyon (Brachypodium) is a wild grass that possesses many features desired in a model plant species, including a small nuclear genome that is less than half the size of a single average wheat chromosome arm. Brachypodium promises to be a potent system for unraveling diverse aspects of grass crop biology. The picture on the cover depicts a spike of the reference Brachypodium inbred line Bd21, whose genome is being sequenced, next to a wheat spike. See Garvin et al. page S-69. Photo by David F. Garvin, USDA-ARS.